火箭浣熊(布莱德利·库珀 Bradley Cooper 饰)偷走了大祭司阿耶莎(伊丽莎白·德比齐 Elizabeth Debicki 饰)的能量电池,包括星爵(克里斯·帕拉特 Chris Pratt 饰)、卡魔拉(佐伊·索尔达娜 Zoe Saldana 饰)、德拉克斯(戴夫·巴蒂斯塔 Dave Bautista 饰)在内的一行人遭到了后者派出的舰队的袭击。一个神秘人物乘坐着飞船救下了银河护卫队的众人,而驾驶着飞船的不是别人,竟然正是星爵的亲生父亲伊戈(库尔特·拉塞尔 Kurt Russell 饰)。
在伊戈的带领下,星爵、卡魔拉和德拉克斯来到了由伊戈一手制造的星球上,而剩下的成员们则着手修复损坏的飞船。那边厢,阿耶莎委托勇度(迈克尔·鲁克 Michael Rooker 饰)抓住星爵替自己报仇,而卡魔拉亦渐渐发现了伊戈和善面目下隐藏的巨大阴谋。
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